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I can't ride a stationary bike for more than 5 minutes without my knees hurting yet I go mountain biking and BMX riding for hours and feel great afterwards. Go figure.
I think when you are running in real life you switch up your gate and stride, you turn, and you speed up and slow down, which is actually better for your joints. With a treadmill, you get stuck in one motion and if that is putting a strain on anything, it really manifests into something.
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