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It could be the way you're running on the treadmill...If it's going too slow so you're running oddly in an unnatural way it can put more pressure on your joints etc...
Maybe up the speed a little? Try doing HIT cardio, sprints for a couple of mins down to slow jogs letting you catch your breath back, then ramped up to sprinting again. Can get a lot more work on the heart and legs with that than running steadily for an hour or so...
Could be a change of shoes needed too, got to have supporting shoes whether on concrete or a treadmill. Can really make a difference.
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