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Over optimization is what always been bad optimization: things that look unnatural.
Having 1000s of "dofollow" links and no "nofollow" links: unnatural.
Having a bunch of links from .gov domains to your penis pill site: unnatural.
Having 1000s of links, but no one ever mentions your site on twitter: unnatural.
Having 100000 links from forum profiles while the other sites in the top 10 for your kw have maybe 200 links in total each: unnatural.
etc.
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